Marco was an undergraduate student in biology in Perugia (Italy) and Oxford (UK) and did his PhD in evolutionary genetics in Pavia (Italy) and Fribourg (Switzerland). He was then a Junior Research Fellow at St. John's College, Oxford, and did research in game theory at Harvard (USA) and in biomedicine in Basel (Switzerland).


  • Applied Game Theory
  • Evolutionary Game Theory
  • Game Theory of Cancer
  • Evolutionary Biology

  • WebSite:

PhD Positions

Feel free to email Marco to discuss projects.

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